While wingless, he often flew on the backs of winged ants under his nefarious mind control.
His next costume had wings, but he pivoted -- strangely enough -- to yellowjackets. Of course, by then he was mentally unstable.
As for dating, he had a queen he was faithful to. A wasp, technically, and she always had wings, at least when she was small. However, he wasn't very good to her.
Somehow, this scene didn't make it into any of the movies:
futurebird